Most leaves that are adjacent in the bound book were not so in the gnawed pile. The asymmetry of the holes shown here (greater for earlier signatures in the alphabet) was created as the worm turned.
News from the Bodleian Libraries Centre for the Study of the Book
Most leaves that are adjacent in the bound book were not so in the gnawed pile. The asymmetry of the holes shown here (greater for earlier signatures in the alphabet) was created as the worm turned.